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Overlapping compartments in non-layout SBML upload

When uploading a pure SBML (no layout), compartments generated by MINERVA overlap.

https://minerva-dev.lcsb.uni.lu/minerva/index.xhtml?id=myoinf_sbml_2

Previously this was not the case (Minerva 12.1.8).

https://curation-sandbox.uni.lu/minerva/index.xhtml?id=mol_targets_SBML2

Attached is the source SBML.

mol_targets_SBML.xml.zip

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Overlapping compartments in non-layout SBML upload